Classification and Systematics of the Ichneumonidae (Hymenoptera) David Wahl American Entomological Institute

ANOMALONINAE

1. Tribe Anomalonini Viereck, 1918 (1900) [= Anomalini of Townes]
   • [Trachynotoidae Förster, 1869. Invalid proposal based on Trachynotus Gravenhorst, junior homonym of Trachynotus Latreille.]
   • Nototrachini Ashmead, 1900. Type-genus: Nototrachys Marshall (= Anomalon Panzer).
   • Anomaloninae Viereck, 1918. Type-genus: Anomalon Panzer. ? correctly formed stem. Names based on Anomalon rather than Nototrachys have been in general use since 1918 and Ashmead's name is therefore not given priority (Article 40 (b) of the Code).
   • Anomalina Viereck; Townes, 1945. This form, with an incorrect stem, is a junior homonym of Anomalitae Blanchard, 1851.
2. Tribe Gravenhorstiini Enderlein, 1912 [includes Podogastrini and Theriini of Townes; Gauld (1976)]
   • [Anomaloidae Förster, 1869. Type-genus: Anomalon Panzer. Förster misidentified the type-genus, as did Ashmead. This error was corrected by Viereck (1918). Förster's name is a junior homonym of Anomalitae Blanchard, 1851. Incorrectly formed stem.]
   • [Anomalonini Ashmead, 1895. ? correctly formed stem. Misidentified type-genus.]
   • [Pharsaliinae Szépligeti, 1905. Invalid proposal based on Pharsalia Cresson, junior homonym of Pharsalia Thomson.]
   • Gravenhorstiini Enderlein, 1912. Type-genus: Gravenhorstia Boie.
   • Hymenopharsalinae Viereck, 1918. Type-genus: Hymenopharsalia Morley (= Ophionellus Westwood). Incorrectly formed stem.
   • Therioninae Viereck, 1918. Type-genus: Therion Curtis. Incorrectly formed stem.
   • Ophionellini Cushman, 1922. Type-genus: Ophionellus Westwood.
   • [Aphanistina Townes, 1944. Not available under Article 13 of the Code.]
   • [Ophiopterina Townes, 1944. Not available under Article 13 of the Code.]
   • [Gravenhorstina Hellén, 1950. Not available under Article 13 of the Code.]
   • [Trichommina Hellén, 1950. Not available under Article 13 of the Code.]
   • [Gravenhorstiina Townes, 1951. Not available under Article 13 of the Code.]
   • Erigorgina Viktorov, 1968. Type-genus: Erigorgus Förster.
   • Podogastrini Townes, 1971. Type-genus: Podogaster Brullé.
   • Theriini Viereck; Townes, 1971. Justified emendation.

1. Tribe Anomalonini
   1. Anomalon Panzer, 1804; worldwide

      Trachynotus Gravenhorst, 1829; preocc. by Latreille, 1829

      Ochlerus Gistel, 1848; preocc. by Spinola, 1837

      Nototrachys Marshall, 1872

      Ophiononeura Cameron, 1904

      Stictophion Cameron, 1906

      Erythrophion Cameron, 1906

      Anomalum Schulz, 1906

      Neogreeneia Viereck, 1912 (Gauld & Bradshaw, 1997)

      Trachyopterus Morley, 1912

      Pseudonototrachys Meyer, 1930

      Microcremastus Hedwig, 1961

2. Tribe Gravenhorstiini
   1. Agrypon Förster, 1860; worldwide

      Agrypum Schulz, 1906

      Trichonotus Cameron, 1906; preocc. by Schneider, 1801, by Rafinesque,1815, and Loew, 1857 (Gauld, 1976a)

      Trichionotus Cameron, 1905 (Gauld, 1976a)

      Odontagrypon Cameron, 1906

      Paragrypon Uchida, 1941 (Gauld, 1976a)

      Dioborus Rao, 1953 (Gauld, 1976a)

   2. Aphanistes Förster, 1869; worldwide except Ethiopian

      Anochilacrum Enderlein, 1921

   3. Atrometoides Fahringer, 1922; Palearctic

      Romanella Meyer, 1935

   4. Atrometus Förster, 1869; Palearctic
   5. Aubertiana Viktorov, 1970; Palearctic

      Aubertia Viktorov, 1968; preocc. by Oberthuer, 1896

   6. Barylypa Förster, 1869; Australian, Holarctic

      Laphyctes Förster, 1869; preocc. by Dujardin, 1844, by Reichenbach, 1850 and Stål, 1853

      Sarntheinia Dalla Torre, 1901

      Hadromanus Szépligeti, 1905 (Gauld, 1976a)

      Magnibucca Morley, 1913

      Trochiscomerus Meyer, 1931 (Gauld, 1976a)

   7. Bimentum Townes, 1971; Ethiopian
   8. Brachynervus Uchida, 1955; Oriental, Palearctic
   9. Calcaneum Townes, 1971; Nearctic
   10. Castrosion Gauld & Bradshaw, 1997; Neotropical
   11. Cechenodes Townes, 1971; Ethiopian
   12. Clatha Cameron, 1905; Oriental
   13. Clypeocampulum Gauld, 1976; Palearctic
   14. Corsoncus Townes, 1971; Neotropical
   15. Encardia Tosquinet, 1896; Ethiopian

       Herus Tosquinet, 1903; preocc. by Rehn, 1900

       Ctenotoma Cameron, 1906

       Ctenocaloides Fahringer, 1936

   16. Gravenhorstia Boie, 1856
       1. subgenus Erigorgus Förster, 1869; Holarctic, Neotropical, Oriental (Gauld, 1976a; Dasch, 1984)

          Sympratis Förster, 1869

          Paranomalon Viereck, 1912

       2. subgenus Gravenhorstia Boie, 1856; Palearctic

          Odontopsis Förster, 1869

       3. subgenus Kokujewiella Shestakov, 1926; Palearctic (Gauld, 1976a)

          Nenethes Ceballos, 1957 (Gauld, 1976a; Atanasov, 1982)

       4. subgenus Ribasia Ceballos, 1921; Palearctic (Gauld, 1976a; Atanasov, 1982)
   17. Habronyx Förster, 1869
       1. subgenus Austranomalon Gauld, 1976a; Australian
       2. subgenus Camposcopus Förster, 1869; Holarctic, Oriental, Australian (Gauld, 1976a)

          Labrorhychus Förster, 1869

          Blaptocampus Thomson, 1892

       3. subgenus Habrocampulum Gauld, 1976; Palearctic (Schnee, 1989)
       4. subgenus Habronyx Förster, 1869; Australian, Neotropical, Oriental, Palearctic

          Acanthostoma Kriechbaumer, 1894

          Macrostemma Shestakov, 1923

          Formosanomalon Uchida, 1928

   18. Heteropelma Wesmael, 1849; worldwide except Ethiopian

       Schizoloma Wesmael, 1849

       Schizopoma Förster, 1869

       Tanypelma Townes, 1971 (Gauld, 1976a)

   19. Indagrypon Nikam, 1982; Oriental
   20. Liopterna Townes, 1971; Nearctic
   21. Metoa Townes, 1971; Nearctic
   22. Neohabronyx Dasch, 1984; Nearctic
   23. Ophionellus Westwood, 1874; Nearctic, Neotropical

       Pharsalia Cresson, 1872; preocc. by Thomson, 1864

       Parophionellus Brues & Richardson, 1913

       Hymenopharsalia Morley, 1913

   24. Ophiopterus Brullé, 1846; Nearctic, Neotropical

       Ophionopterus Schulz, 1906

       Ophionopterus Morley, 1912

   25. Parania Morley, 1913; worldwide except Australian
   26. Perisphincter Townes, 1961b; Australian, Neotropical, Oriental
   27. Phaenolabrorychus Viereck, 1913; Neotropical
   28. Philodrymus Townes, 1971; Neotropical
   29. Podogaster Brullé, 1846; Neotropical
   30. Porizonopteron Meyer, 1931; Palearctic
   31. Pseudagrypon Lee & Kim, 1984; Palearctic
   32. Pseudanomalon Szépligeti, 1905; Ethiopian, Oriental
   33. Sphaeromanus Aubert, 1979; Palearctic
   34. Spolas Townes, 1961b; Hawaii, New Zealand
   35. Stangepelma Porter, 1976; Neotropical
   36. Therion Curtis, 1829-30; Holarctic, Neotropical, Oriental

       Therium Agassiz, 1846

       Exochilum Wesmael, 1849

   37. Trichomma Wesmael, 1849; worldwide

       Trichomella Szépligeti, 1910

   38. Vernamalon Gauld, 1976a; Ethiopian