Monophyly of Hemistephanus
The following synapomorphies support Hemistephanus as a genus distinct from Megischus, contrary to the opinion of Townes2 and Gauld11.

Synapomorphies for Hemistephanus
Qualitative and Biometric Analysis
80 characters (15 biometric) were proposed as a new character-set for the group. A hierarchical list of biometrical ratios is presented on table 1.
Influence of Size and Sex on the Variation of Characters
Smaller specimens often have more distortion or masking of qualitative and biometric characters than larger ones. Thus, for a given species, an unusually small specimen is generally much less characteristic than a very large one.
The morphometric variation is considerably different for males and females. This can be more easily observed with the ratio Pnl/Hd, which has low taxonomic value for females, but is the most accurate for males (Table 1).
| Table 1. Ten more important ratios, ranked in descending order of taxonomic importance, for females and males of Hemistephanus. Relative importance of ratio Pnl/Hd indicated by a pointer. | ||
| Females | Males | |
| 01. Ov/Pt | 01. Pnl/Hd ![]() |
|
| 02. Ov/Tt | 02. io/oc | |
| 03. io/oo | 03. Pnc/Pnl | |
| 04. Pnc/Pnl | 04. io/io | |
| 05. io/oc | 05. 2-1A/1Cu | |
| 06. btc/btw | 06. Pt/Gs | |
| 07. 2-1A/1Cu | 07. btc/btw | |
| 08. fl2/fl1 | 08. fl2/fl1 | |
| 09. Pt/Gs | 09. Pt/Ptw | |
10. Pnl/Hd ![]() | 10. T4c/T4l | |
Species Classification
The dendrogram below proposes a classification for the species. The numbers correspond to the species number shown on the graphs with morphometric data (see M&M).
Classification of Hemistephanus species (not a cladogram)
Geographical Distribution
The geographical distribution of the genus Hemistephanus is presented below. This genus does not occur in Australia, as previously suggested 12,15,16.

This page last updated: April 25, 1998.