
Classification of Extant Hymenoptera (based on Naumann 1991)
The generally used classification of Hymenoptera does not yet reflect our knowledge of the phylogeny of the order. Traditionally, two suborders are recognized: Symphyta (paraphyletic or perhaps even polyphyletic) and Apocrita. Within the Apocrita the Aculeata form a monophyletic group. In the older literature and even today for convenience, the remaining superfamilies may grouped together as the Terebrantes or Parasitica. Rasnitsyn (1988) has proposed an alternative classification, incorporating extinct taxa, that may be viewed here.
Order Hymenoptera
Suborder Symphyta
Superfamily Xyeloidea
Superfamily Megalodontoidea
Superfamily Cephoidea
Superfamily Siricoidea
Superfamily Tenthredinoidea
Suborder Apocrita
Superfamily Stephanoidea
Superfamily Trgonalyoidea
Superfamily Megalyroidea
Superfamily Ceraphronoidea
Superfamily Evanioidea
Superfamily Ichneumonoidea
Superfamily Proctotrupoidea
Superfamily Platygastroidea
Superfamily Cynipoidea
Superfamily Chalcidoidea
"Aculeata"
Superfamily Chrysidoidea
Superfamily Vespoidea
Superfamily Sphecoidea
Superfamily Apoidea
,==Cimbicidae
,======|
| `==Tenthredinidae
|
,===| ,==Argidae
| | ,==|
| | | `==Pergidae
| `===|
,==| |
| | `=====Electrotomidae
,===| |
| | `=============Blasticotomidae
,==| |
| | `================Xyelotomidae
| |
| `====================Xyelidae: Macroxyelinae
|
<<===|
| ,================Xyelidae: Xyelinae
| |
| ,===| ,============Xyelydidae
| | | |
| | `===| ,========Pamphiliidae
| | | |
| | `===|
| | | ,||||Praesiricidae
| | `===|
`==| `====Megalodontidae
|
| ,========Sepulcidae
| ,========|
| | `========Cephidae
| |
`==| ,========Anaxyelidae
| ,=====|
| | `========Gigasiricidae
`==|
| ,===========Siricidae
| |
`==| ,========Xiphydriidae
| |
`==| ,||Paroryssidae
| ,==|
| | `==Orussidae
`==|
`=====Apocrita
Containing clade: Endopterygota
Sources of Information on the Internet
References
Boucek, Z. 1988. Australasian Chalcidoidea (Hymenoptera): a biosystematic
revision of genera of fourteen families, with a reclassification
of species. C.A.B. International, Oxon.
Brothers, D.J. 1975. Phylogeny and classification of the aculeate
Hymenoptera, with special reference to Mutillidae.
Kans. Univ. Sci. Bull. 50:483-648.
Carpenter, J. M. 1987. On the evolutionary genetics of social
wasps and the phylogeny of the Vespinae (Hymenoptera,
Vespidae). Insectes Sociaux 34(1): 58-64.
Crozier, R. H. 1975. Hymenoptera. Borntraeger, Berlin; Stuttgart.
Danforth, B. D. 1989. The evolution of hymenopteran wings: The
importance of size. Journal Of Zoology 218(2): 247-276.
Fergusson, N. D. M. 1988. A comparative study of the structures
of phylogenetic importance of female genitalia of the
Cynipoidea (Hymenoptera). Systematic Entomology 13(1):
13-30.
Gauld, I. and Bolton, B. 1988. The Hymenoptera. British Museum
(Natural History); Oxford University Press, London;
Oxford; New York.
Gibson, G.A.P. 1985. Some pro- and mesothoracic structures
important for phylogenetic analysis of Hymenoptera,
with a review of terms used for the structures. Can.
Ent. 117:1395-1443.
Gibson, G.A.P. 1986. Evidence for the monophyly and relationships
of Chalcidoidea, Mymaridae, and Mymarommatidae (Hymenoptera:
Terebrantes). Can. Ent. 118:205-240.
Johnson, N. F. 1988. Midcoxal articulations and the phylogeny
of the order Hymenoptera. Annals Of The Entomological
Society Of America 81(6): 870-881.
Krombein, K. V. 1979. Catalog of Hymenoptera in America north
of Mexico. Smithsonian Institution Press, Washington.
Leluk, J., J. Schmidt, et al. 1989. Comparative studies on the
protein composition of hymenopteran venom reservoirs.
Toxicon 27(1): 105-114.
Plant, J. D. and H. F. Paulus. 1987. Comparative morphology of
the postmentum of bees (Hymenoptera: Apoidea) with
special remarks on the evolution of the lorum. Zeitschrift
Fuer Zoologische Systematik Und Evolutionsforschung
25(2): 81-103.
Quicke, D. L. J., M. G. Fitton, et al. 1992. Phylogenetic implications
of the structure and distribution of ovipositor valvilli
in the Hymenoptera (Insecta). Journal Of Natural History
26(3): 587-608.
Quicke, D. L. J., S. N. Ingram, et al. 1992. Sperm structure
and ultrastructure in the Hymenoptera (Insecta). Zoologica
Scripta 21(4): 381-402.
Rasnitsyn, A. P. 1988. An outline of evolution of the hymenopterous
insects (Order Vespina). Oriental Insects 22: 115-145.
Saini, M. S. 1986. Comparative studies on the metapostnotal modifications
in suborder Symphyta and Hymenoptera Parasitica. Journal
Of Entomological Research 10(1): 85-90.
Schmidt, J. O., M. S. Blum, et al. 1986. Comparative enzymology
of venoms from stinging Hymenoptera. Toxicon 24(9):
907-922.
Schoenitzer, K. and G. Lawitzky. 1987. A phylogenetic study of
the antenna cleaner in Formicidae, Mutillidae, and
Tiphiidae (Insecta, Hymenoptera). Zoomorphology 107(5):
273-286.
Whitfield, J. B., N. F. Johnson, et al. 1989. Identity and phylogenetic
significance of the metapostnotum in nonaculeate Hymenoptera.
Annals Of The Entomological Society Of America 82(6):
663-673.
![[Top of Page]](/gifs/newtools/up.arrow.gif)